Core Asterids Diversity and Evolution of Asterids viburnums umbels and lobelias will examine the second of the two well supported lineages of the core asterids lamiids campanulids lamiid or Asterid I group campanulid or Asterid II group appears to have the fused corolla derived independently via a different floral developmental pathway relative to the Asterid I group Lecture exam 2 mean 76 2 80 range 61 85 Lab exam 2 mean 37 7 45 range 21 45 Early vs Late Sympetaly euasterids II campanulids Dipsacales euasterids I lamiids order within campanulids or Asterid II group lamiids 2 or 7 families and nearly 1 100 species of herbs and shrubs largely from North Hemisphere iridoids opposite leaves inferior gynoecium campanulids inferior G Calendula Asteraceae early Anchusa Boraginaceae late Caprifoliaceae what is it DNA phylogeny Adoxa pre DNA era four families were recognized as in Gleason Cronquist Caprifoliaceae what is it DNA phylogeny Adoxa will use 2 here honeysuckle honeysuckle Dipsacaceae Valerianaceae teasel valerian Caprifoliaceae what is it DNA phylogeny elderberry post DNA era up to 7 families can be recognized Caprifoliaceae viburnum viburnum teasel valerian Caprifoliaceae what is it the two families can be differentiated by flowers the two families can be differentiated by flowers short styled Adoxaceae long styled Caprifoliaceae short styled Adoxaceae long styled Caprifoliaceae Viburnum viburnum Lonicera honeysuckle Adoxaceae viburnums Adoxaceae viburnums 5 genera and 200 species of opposite leaved shrubs and some herbs mainly North Temperate flowers small and usually in flat topped cymes 5 merous with up to 5 fused carpels and short style berry or drupaceous fruits Sambucus racemosa red berried elder elder berries are opposite compounded leaved shrubs Sambucus canadensis Common elder Adoxaceae viburnums viburnums are simple leaved shrubs but often lobed or coarsely toothed Adoxaceae viburnums note enlarged sterile flowers on edge of head inflorescence for pollinator attraction Viburnum opulus trilobum high bush cranberry Viburnum davidii Adoxaceae viburnums many viburnums have glands on petioles Adoxaceae viburnums Adoxa moschatellina muskroot Viburnum lentago nannyberry unusual reduced woodland herb with musky scent and trifoliate spiralled leaves Viburnum acerifoliummaple leaved viburnum Viburnum rafinesquianum arrowwood Caprifoliaceae honeysuckles Northern hemisphere family and tropical mountains of 43 genera and about 900 species of shrubs subshrubs vines and herbs Caprifoliaceae honeysuckles CA 5 CO 5 A 4 5 G 2 5 5 merous long styled and bell shaped or strongly zygomorphic inferior ovary forms berry or reduced to achenes Lonicera honeysuckle Linnaea twinflower Lonicera honeysuckle Dipsacus teasel Triosteum horse gentian Caprifoliaceae honeysuckles Caprifoliaceae honeysuckles honeysuckles include native right and introduced aggressive shrubs below or vines natives Lonicera dioica red honeysuckle Lonicera tatarica tartarian honeysuckle Lonicera canadensis fly honeysuckle Caprifoliaceae honeysuckles Lonicera reticulata grape honeysuckle Caprifoliaceae honeysuckles invasives Lonicera japonica Diervilla lonicera bush honeysuckle Lonicera xylosteum European fly honeysuckle native leaf miners Lonicera X bella pretty honeysuckle Triosteum perfoliatum Horse gentian Caprifoliaceae honeysuckles Linnaea borealis twinflower Caprifoliaceae honeysuckles many culativated ornamentals Characteristic subshrub of circum boreal forests where it forms large colonies Two flowers form per inflorescence Abelia Caprifoliaceae honeysuckles Kolkwitzia Caprifoliaceae honeysuckles the old Valerianaceae and Dipsacaceae exhibit features that will be seen in Asteraceae the old Valerianaceae and Dipsacaceae exhibit features that will be seen in Asteraceae congested inflorescences bracted inflorescences reduced calyx pappus one seeded achene Knautia pappus Valeriana uliginosa marsh valerian Valeriana edulis edible valerian Knautia Caprifoliaceae honeysuckles Apiales the old Valerianaceae and Dipsacaceae exhibit features that will be seen in Asteraceae involucral bracts 7 families and nearly 5 500 species of herbs and shrubs Australasia appears to be center of diversity iridoids aromatic alternate compound leaves umbels corolla separate inferior gynoecium lamiids campanulids inferior G Dipsacus fullonum teasel Introduced and adventive Involucral bracts ginseng Araliaceae pappus Araliaceae ginseng tropical family of trees shrubs or herbs temperate leaves divided or up to 3 times or more compound distinctive aroma and chemicals parsnip Apiaceae Araliaceae ginseng CA 5 CO 5 A 5 G 5 umbels flowers small non asterid like no corolla tube 5 merous with 5 fused carpels forming inferior ovary berries Aralia hispida bristly sarsaparilla note 5 styles Aralia hispida bristly sarsaparilla Aralia nudicaulis wild sarsaparilla Araliaceae ginseng Aralia elata Hercule s club Devil s walking stick Araliaceae ginseng Panax quinquefolius American ginseng Rich woods leaflets are stalked Heavily collected woodland species for roots Aralia racemosa spikenard Araliaceae ginseng Panax trifolius Dwarf ginseng Rich woods leaflets are sessile Araliaceae ginseng Hedera helix English ivy Eurasia N Africa invasive in U S Araliaceae ginseng Tetrapanax papyrifera rice paper Schefflera umbrella plants Apiaceae umbels aromatic herbs with hollow stems dissected or compound leaves leaves strongly sheathing Apiaceae umbels Large family of 430 genera and over 3700 species most common in north temperate regions Economically important carrot parsnip parsley celery dill caraway parsnip carrot Apiaceae umbels CA 5 CO 5 A 5 G 2 flowers small in umbels often compound female flowers often along edge of each umbellet common feature in Asterids with the formation of a head sheath Heracleum lanatum cow parsnip Apiaceae umbels Apiaceae umbels CA 5 CO 5 A 5 G 2 CA 5 CO 5 A 5 G 2 flowers small in umbels often compound female flowers often along edge of each umbellet 5 merous with no corolla tube inferior gynoecium of 2 carpels separating at maturity fruit dehiscent and splitting schizocarp 2 dry 1 seeded mericarps held together by carpophore fruits 5 ribbed separated by oil canals taxonomic character for separating genera Apiaceae umbels Apiaceae umbels Cicuta bulbifera Bulblet water hemlock Cicuta maculata spotted water hemlock Angelica atropurpurea great angelica Large coarse herb of wetter areas Although containing nasty compounds Cicuta is not the