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evolution
a change in allele frequencies in a population over time
scientific knowledge
empirical & testable
scientific theory
well-established explanation of some part of the world that can incorporate facts, laws, inferences, and tested hypothesis; includes multiple parts
gene
small section of DNA that codes for a protein
alleles
variants of a gene
microevolution
genetic changes within a species; short time frame
macroevolution
origin and extinction of species; long time frame   
european thought during middle ages
1. earth young & at center of universe 2. fixity of species (no evolution) 3. humans at top of scala naturae 4. organisms successful at what they do, because God designed them that way
james ussher & john lightfoot
james believed earth was created sunday, oct 23, 4004 bc; john added at 9 am london time
plato
world of ideals - for everything, there is an ideal or perfect counterpart
aristotle
scala naturae (ladder of nature) - organisms can be grouped on a linear scale from simple to complex
teleology
use of ultimate purpose or design as a means of explaining natural phenomena
copernicus
made observations of night sky & discovered earth is not at the center of the universe
buffon
french aristocrat & naturalist - historie natuelle 1. earth > 6000 yrs old 2. species change overtime 3. noted anotomical similarity btwn humans & apes & discussed possibility of common ancestry; no mechanism, though
jean baptiste lamarck
philosophie zoologique; 1st to propose a coherent mechanism to explain how evolution may occur; acquired characteristics passed onto offspring (WRONG)
thomas malthus
essay on the principle of population; compared european pop growth with food production capacity (pop growth faster); if left unchecked, pop will tend to increase faster than food resources
new ideas around darwin's time
1. world not at center of universe 2. humans are animals & have same basic anatomical features as many other animals 3. species can change over long pds of time in response to changes in environment 4. some species alive in past were not alive today 5. incremental changes over long pd…
4 mechanisms of evolution
natural selection mutation genetic drift gene flow
modern synthesis
natural selection & other evolution mechanisms + laws of inheritability + understanding of protein synthesis
biological species
group of organisms that are 1. actually or potentially interbreeding and 2. reproductively isolated from other groups
reproductive isolation
something that prevents act of mating / production of viable offspring rather than physical separation; parts don't fit, diff body sizes, diff courtship rituals, genetical differences cause offspring to be sterile (mule)
2 examples of exceptions to reproductive isolation
male tiger & female lion; some guenon species
3 modes of speciation
allopatric, parapatric, & sympatric
allopatric speciation
pop is divided by a physical barrier (mountain, river) & then separated populations diverge over time
parapatric evolution
2 pops are physically adjacent but adapt to diff environmental conditions over time; hybrid zones occur btwn diff species ranges
sympatric speciation
indivs of a single pop in 1 habitat diverge over time; no large-scale separation or barrier; rarest mode of speciation; ex: tree hopper
cladogenesis
speciation by splitting of lineages
adaptive radiation
special type of cladogenesis; rapid series of multiple cladogenic events; often occurs following colonization of a new landmass; ex: hawaiian honeycreepers, lemurs
anagenesis
gradual transformation of one species into another; deciding where to recognize a species boundary is a problem
adaption
any trait that increases an organism's fitness (lifetime reproductive potential); must be maintained by natural selection
exaptation
when an adaption evolves for one function, but is then useful for another; ex: feathers - first for warmth in dinosaurs, then for flight in birds
taxonomy
scientific classification, usually based on shared characteristics
phylogeny
model of presumed evolutionary relationships based on shared characteristics
primitive characters
inherited from a more distant common ancestor; plesiomorphy
derived characters
evolutionary innovations that first appeared in the common ancestor of the group; apomorphy
convergence
similar function, but diff body parts; ex: flying lemur, gliding lizard, gliding snake
parallelism
similar function, same structures being used; more recent common ancestry that convergence; ex: parallel evolution of prehensile tails in S. America
evolutionary reversal
re-evolution of a plesiomorphic feature
2 rules for reconstructing phylogenies
1. you can't use symplesiomorphies to recognize clades, only synapomorphies 2. homoplasy is common & has the potential to mess up your phylogenetic reconstruction
principle of parsimony
parsimony = stinginess; assumes that features will evolve as few times as possible; minimizes homoplasy. these rules = cladistic methodology
retrotransposons
sequences of DNA that can copy themselves at random & reinsert themselves in the genome at random
chicxulub impact crater
asteroid that ended dinosaur pd; may be 3rd largest crater on earth; 200 km diameter; marks boundary between cretaceous & paleogene periods
galileo
refined telescope lens & looked at night sky; confirmed copernicus is right
carolus linnaeus
founded modern taxonomy with system nature; grouped humans with other primates
richard owen
anti-evolutionist; homology: same origin in diff animals under every variety of form & function; there is an archetype for living organisms/common anatomical plans for diff groups
georges culvier
anti-evolutionist & critic of lamarck; worked with fossils from paris basin; established extinction as a fact; catastrophism provided evidence that earth was old
charles lyell
scottish geologist; uniformitarianism; given enough time small changes can have a large effect; earth is very old
charles darwin
origin of species; natural selection
3 fundamental points of natural selection
1. ability of pop to expand is infinite but carrying capacity of environment finite 2. indivs can vary in morphology & behavior, differential surviving & reproduction 3. some of this variation is transmitted from parents to offspring
for natural selection to occur...
1. trait must be heritable 2. must be variation 3. must be differential reproduction
fitness
relative measure of lifetime reproductive success
types of mutagens
radiation, chemicals, viruses
eocene epoch
warmest period of time in last 65.5 million years; widespread tropical forests
blending inheritance
a model of inheritance in which the hereditary material of the mother and father was thought to combine irreversibly in the offspring
genotype
genetic makeup
phenotype
observable traits
mutation
random changes in the DNA sequence; ultimate source of new genetic variation in a population
genetic drift
random change in allele frequencies within a population that is produced by the random factors of inheritance and population subdivision
gene flow
the movement of alleles btwn populations through both migration and interbreeding of indivs from diff pops
biological species
a group of organisms that can't interbreed in nature
ecological species
absence of interbreeding btwn two pops is not a necessary condition for defining them as separate species
3 postulates of natural selection
limited resources, variation, & inheritance
homologous character
features that are shared by 2 organisms because they are inherited from a common ancestor
analogous characters
similar anatomical structures that evolved independently to accommodate a similar functional demand
homoplasy
similarity between 2+ taxa that is not inherited from their common ancestor; false similarity; usually in response to similar niche/environmental pressures; 3 types: convergence, parallelism, & evolutionary reversal
symplesiomorphy
shared primitive characters
synapomorphy
shared derived character
axis of rotation
point around which the lever moves
force arm
perpendicular distance btwn AOR & line of action of an applied force
load (resistance) arm
perpendicular distance btwn AOR & line of action of a load
1st, 2nd, & 3rd class lever
1: A 2: R- power 3: F- speed

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