Low rate of lineage diversification High rates of lineage diversification Ancestral trait innovation Evolutionary dead ends (e.g. specialization hypothesis) Key innovation hypothesis for diversity Low rate of trait diversification no special name? Non-adaptive radiation High rate of trait diversification Adaptive divergence? Local adaptation? Adaptive radiation Niche conservatismHarmon et al. 2003 ScienceHarmon 03-3 Harmon et al. 2003 Science early diversification -> greater phylogenetic signalMeasuring niche conservatism - phylogenetic signal!K: Blomberg et al. (2003) Evolution; examples: Ackerly, PNAS in review!Blomberg’s K: measures degree of similarity among close relatives, relative to expectations based on Brownian motion K<<1 K~1 K>>1 convergence brownian
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