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anthron pous/podos
joint foot
most diverse phlyum, 3/4 of all known species of animals, occur in all habitats chitinous, segmented exoskeleton (protective while flexible) segmentation - efficient air piped directly to cells- high metabolic rate highly developed sensory complex behavior patterns
Phylum Anthropoda
evolved before Cambrian, extinct for 200 million years; abundant during Cambrian and Ordovician dorsally flattened bottom dwellers; scavengers chitinous exoskeleton (calcium carbonate) tagmata: head, trunk, pygidium cephalon with antennae, compound eyes, mouth, four pairs of legs no …
Subphylum Trilobita
giant water scorpions (extinct) head with six fused segments, simple eyes, chelicerae and pedipalps
Class Merostomata, Subclass Eurypterida
horseshoe crabs, ancient, Cambrian four species in three genera extant unsegmented, carapace, broad abdomen, long tail spike (telson) chelicerae, one pair of pedipalms, four pairs of legs book gills - tracheal system that goes to book gills two lateral eyes and two simple eyes feed …
Subclass Xiphosurida
sea spiders; 100 species four pairs of legs some duplicated segments male extra legs called ovigers circulatory system simple; excretory and resp system - absent digestive system and gonads branch into legs
Class Pycnogonida
spiders, scorpions, pseudoscorpions, whip scorpions, ticks, mites, daddy longlegs 80,000 species - first terrestrial two tagmata: cephalothorax and abdomen chelicerae modified into fangs, claws into pedipalps
Class Arachnida
40,000 species cephalothorax called prosoma, abdomen- opisthosoma - unsegmented chelicerae, terminal fangs leglike pedipalps- transfer sperm four pairs of legs
Order Araneae
breathe through book lungs, trachea evolved indpendently of insects malpighian tubules eight simple eyes - vision usually poor mechanoreceptors on body, sensory setae
Spider organs and senses
courtship ritual precedes mating use pedipals for depositing sperm and into genital opening lay egg in silken net
Spider reproduction
scorpions; 1400 species; ancient of arachnids predatory mostly on insects and spiders sieze with pedipalps and shred with chelicerae short cephathor, pedipalps, walking legs, one pair of median eyes, up to 5 pairs of lateral eyes metasoma of five segments with stinger comblike pectin…
Order Scorpiones
complex mating dance, hold female chelae as steps back and forth sometimes stings her deposits on substrate brood and nourish young in female's reproductive tract babies crawl onto mom's back until first molt
Scorpion reproduction
sun/camel spiders nonvenomous, shred prey with chelicerae, up to 15 cm across; desert habitats; high metabolism
Order Solpugida
daddy longless, 5000 species, harvestment abdomen and cephthor round and broad no pedicel external segmentation four pairs of legs carnivorous and scavengers nonvenomous
Order Opiliones
ticks and mites 40,000 species most habitats; eyelashes/dust mites
Order Acari
complete fusion of cephalothorax and abdomen, no division mouth parts called capitulum chelicera pedipalps form hypostome; rostrum/tectum extends dorsally four pairs of legs spermatophores larvae with six legs; some eight legged
Acarian anatomy
'many footed'; two tagmata: head and trunk paired appendages trachea carry respiratory gasses to and from all body cells independent from arachnids and insects; third evolution escretion through Malpighian tubules
Subphylum Myriapoda
centipeds (cheilos- margin) carnivorous; some dangerously venomous 177 segments; one pair of jointed legs first segments make venomous claws one pair of antennae, mandibles, one/two maxillae pair of eyes on top- ocelli digestive straight tube respiration through spiracles sexes se…
Class Chilopoda
millipedes slow, graceful some secrete toxic or repellant fluids from repugnatorial glands more than 10,000 species 25-100 segments; two pairs of legs two pairs of spiracles, two genital apertures females lay eggs in nest and guard them larval only have one pair of legs
Class Diplopoda
pauros (small) soft bodied- 500 species small head with branched antennae no true eyes; 12 segments no tracheae, spiracles or circulatory system
Class Pauropoda
small, centipede like bodies pests on crops 14 segments, 12 have legs, antennae long and unbranched eyeless; spiracles on head and tracheal tubes to anterior segments
Class Symphyla
67,000 species two pairs of antennae, one pair of mandible, two pairs of maxillae, pairs of appendages on each segment biramous respiratory organs are gills segments are fused- carapace bodies covered with secreted cuticle (chitin proten, calcium carbonate) soft and thin at joints e…
Phylum Arhtropoda; Subphlyum Crastacea
Serial homology -Protopod: basal portion, two parts - Exopod: laterally borne by protopod, one to several parts - Endopod: medially borne by the protopod, one to several - Exites and endites: respectively lateral and medial - Uropod: tail fin
Appendages of crustaceans
hemocoel; persistent with blastocoel with hemolymph striated muscles in antagonistic groups flexors: draw a part towards the body extensors: extend part outward gas exchange in small crustaceans occur across thin areas of cuticle; larger have internal gills
Crustacean internal features
open system; hemolymp returns to sinuses; valves prevent backflow pigment can be colorless, red or blue dorsal heart the chief organ excretory system is a pair of tubular antennal or maxillary glands- each gland has a vesicle; spongy mass called labyrinth excretion of nitrogenous wast…
Circulatory and excretory system of crustaceans
similar to annelids brain is pair of supraesophageal ganglia eyes have units called ommatidia, covered by cornea, makes a mosaic statocysts contain statoliths; tactile hairs
Nervous and sensory system of crustaceans
separate sexes, lots of specialization for copulation parthenogenetic brood their eggs direct devleopment; some have multiphasic life cycles ancestral nauplius- widely occurring larvae
Reproduction in crustaceans
1)outermost, lipid impregnated protein 2)bulk of cuticle, several layers 3)innermost -intermolt phases are called instars; shedding the cuticle for growth
1)epicuticle 2)procuticle 3)endocuticle: chitinous, uncalcified Molting and ecdysis
-initiated by environmental stimuli - stops production of molt inhibiting hormone from the x-hormone -promotes release of molting hormone from the Y-organs, initiate molting -androgenic glands occur in some males
Hormonal control in crustaceans
- vary greatly, much specialization - predators eat mostly larvae - scavengers eat dead - suspension feeders - crayfish have two part stomach with gastric mill
Crustacean feeding
ten species; all from caves into ocean trunk segments, identical, paired, biramous, swmming appendages angennules biramous, maxillae and maxillipeds prehensile
Class Remipedia
only nine species, live in sediment thoracic limbs all similar no eyes, carapace, or abdominal appendages true hermaphrodites, eggs and sperm through common duct
Class Cephalocarida
10,000 species Three orders: Anostraca (fairy shrimp), Notostraca (tadpole), Diplostraca (water fleas) flattened, leaflike phyllopodia or legs as resp. organs mostly freshwater complex life cycles
Class Branchiopoda
10,000 species five cephalic, six thoracic, four abdominal segments nauplii have maxillopodon eye (many subclasses)
Class Maxillopoda
only ten species; live in interstitial spaces in sand
Subclass Mystacocarida
abundant lack carapace, retain nauplius eye into adulthood, single uniramous maxillipeds, four pairs of swimming appendages, fifth pair of legs reduced most free living but many parasitic direct development; parasitic- multiphasic
Subclass Copepoda
only described in 1983, 12 species tiny one pair of antennae on sexual females parthogenetic and sexual life cycle parasitic- penetrate cuticle of hosts by mouth tube abdomen and thoracic limbs lost during metamorphosis juveniles have six or seven abdominal segments
Subclass Tantulocarida
small group of fish ectoparasites with sucking mouth shieldlike carapace; compound eyes; four biramous throacic appendages second maxillae modified as suction cups no nauplius, resemble adults
Subclass Branchiura
130 species of wormlike parasites- tongueworms lung parasites of vertebrates transverse rings make look segmented nonchitinous, molted during larval stages five protuberances on anterior end simple, digestive system no circulatory, excretory or respiratory sexes separate, females l…
Subclass Pentastomida
barnacles; three smaller orders of burrowing or parasitic forms sessile head reduced, no abdomen, thoracic legs, many cirri with hairlike setae hermaphroditic, internal fertilization, free living cyprid larvae metamorphose into adult longest penis
Subclass Cirripedia
20,000 species; largest class, great diversity
Class Malacostraca
roly polys and friends no carapace, sessile compound eyes maxillipeds are first pair of thoracic limbs, lack exopods direct development; few parasitic forms with complex life cycle
Order Isopoda
like isopods but compressed laterally gills in thoracic position one pair of limbs for swimming and one pair for jumping direct development, no metamorphosis
Order Amphipoda
90 species, krill carapace fused not entirely enclose the gills no maxillipeds, with exopods bioluminescent because of photophore
Order Euphausiacea
18,000 species three pairs of maxillipeds and five pairs of walking legs some legs modify to form chelae (claws) largest arhtropod is Japanese spider crab
Order Decapoda

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