Rutgers University MBB 408 - Lecture 2 - Light Reactions of Photosynthesis (9 pages)

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Lecture 2 - Light Reactions of Photosynthesis



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Lecture 2 - Light Reactions of Photosynthesis

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Pages:
9
School:
Rutgers University- The State University of New Jersey
Course:
Mbb 408 - Molecular Biology & Biochemistry

Unformatted text preview:

694 40 8 115 512 M ol ec ul ar Bi olo gy Bi oc he mi str y Lecture 2 Light Reactions of Photosynthesis November 4 2008 1 Overall photosynthetic process light converted into chemical energy for biosphere provides atmospheric O2 fixes atmospheric CO2 into CH2O 1011 tons worldwide yr 2 Photosynthesis consists of light reactions drive O2 evolution generate NADPH and ATP and carbon assimilation reactions NADPH and ATP drive CO2 fixation via ribulose 1 5 bis P carboxylase Calvin cycle to regenerate ribulose 1 5 bis P Improved understanding of light reactions is basis for harnessing solar energy a sustainable and renewable source of clean energy 3 Photosynthesis occurs in chloroplasts in higher phototrophs light reactions in thylakoid membrane stacked grana and unstacked stroma lamellae regions thylakoid lumen accumulates H CO2 fixation occurs in stroma 4 Distinguishing oxygenic photosynthesis H2O as e donor algae plants and anoxygenic H2S organic acids alcohols as e donors green and purple bacteria 5 Photosynthetic pigments associated noncovalently in pigmentprotein complexes a Chlorophyll Chl Mg tetrapyrrole has light harvesting role funnels collected energy to reaction center and photochemical reaction center role catalyzes charge separation b Carotenoids carotene accessory light harvesting and photoprotective roles 6 Fates of photoexcitations resonance energy transfer in lightharvesting proteins e g light harvesting complex II LHC II electron transfer initiated in reaction center loss as fluorescence useful in energy transfer studies and radiationless decay 7 Molecular architecture and role of LHC II major light harvesting complex of higher plants X ray structure reveals how LHC II Chls funnel excitations to PSII 8 Oxygenic phototrophs use two photosystems PSI and PSII multisubunit integral membrane complexes with reaction center and electron transport components surrounded by light harvesting Chls a PS I reaction center P700 absorbs 700 nm photons strong reductant



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