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Ecology
scientific study of the interactions between organisms and the environment
Organismal Ecology
studies how an organism's structure, physiology and (for animals) behavior meet environmental challenges
Population Ecology
focuses on factors affecting how many individuals of a species live in an area
Community Ecology
deals with the whole array of interacting species in a community
Ecosystem Ecology
emphasizes energy flow and chemical cycling among the various biotic and abiotic components
Landscape
mosaic of connected ecosystems
Landscape Ecology
deals with arrays of ecosystems and how they are arranged in a geographic region
Biosphere
global ecosystem, the sum of all the planet's ecosystems
Global Ecology
examines the influence of energy and materials on organisms across the biosphere
Rachel Casron
credited with starting the modern environmental movement with the publication of 'Silent Spring' in 1962
Biotic
living factors
Abiotic
nonliving factors
Factors that determine distribution
biotic and abiotic
Dispersal
movement of individuals away from centers of high population density of from their area of origin
Community
an assembly of populations of various species living close enough for potential interaction
Interspecific Interactions
relationships between species in a community
Examples of Interspecific interactions
competition, predation, herbivory, and symbiosis 
What does interspecific interactions affect?
can affect the survival and reproduction of each species and the effects can be summarized as positive, negative or no effect
Interspecific Competition 
-/- interaction; occurs when species compete for a resource in short supply
Competitive Exclusion
local elimination of competing species; strong competition can lead to this
Predation 
+/- interaction; interaction where one species, the predator, kills and eats the other, the prey
Feeding adaptations of predators
claws, teeth, fangs, stingers, and poison
Behavioral Defenses
hiding, fleeing, forming herds and schools, self-defense, alarm calls and some morphological and physiological defense adaptations
Cryptic coloration
camouflage; makes prey difficult to spot
Herbivory
+/- interaction; a herbivore eats parts of a plant or alga
Symbosis 
relationship where two or more species live in direct and intimate contact with eachother
3 types of symbiosis
mutualism, commensalism, parastitism 
Mutualism
+/+ interaction; interspecific interaction that benefits both species
Obligate (mutualism)
one species cannot survive without the other
Faculative (mutualism)
where both species can survive alone
Commensalism
+/0 interaction; one species benefits and the other is apparently unaffected
Why is commensal interactions hard to document in nature?
any close association likely affects both species
Parasitism
+/- interaction; when one organism, the parasite, derives nourishment from another organism, its host, which is harmed in the process
Endoparasites
parasites that live within the body of their host
Ectoparasites
parasites that live on the external surface of their host
Species diversity
variety of organisms that make up the community
Trophic structure
feeding relationships between organisms in a community; key factor in community dynamics
Food chain
links trophic levels from producres to top carnivores
Two fundamental features of a community structure
species diversity and trophic structure
Food web
branching food chain with complex trophic interactions
How can food webs be simplified?
isolate a portion of a community that interacts very little with the rest of the community
Can species play a role at more than one trophic level?
Yes
Limits on Food Chain Length
is usually only a few links long
Two hypotheses to explain food chain length
energetic hypothesis and dynamic stability hypothesis
Energetic hypothesis
suggests that the length is limited by inefficient energy transfer
Dynamic stability hypothesis
proposes that long food chain are less stable than short ones
Which hypothsis is most supported?
most data supports the energetic hypothesis
Types of species with a large impact
dominant species, invasive species, keystone species, and foundation species 
Dominant Species
those that are most abundant or have the highest biomass; exert powerful control over occurrence and distribution of other species
Biomass
total mass of all individuals in a population
Hypothesis suggestions on dominant species 
most competitive in exploiting resources; most successful at avoiding predators
Invasive species
typically introduced to a new environment by humans, often lack predators or disease
Keystone Species
exert strong control on a community by their ecological roles, or niches; not necessarily abundant in a community
Foundation species
"ecosystem engineers" cause physical changes in the environment that affect community structure 
Facilitators
some foundation species act as these; have positive effects on survival and reproduction of some other species in the community
Bottom-up Model of community organization
proposes a unidirectional influence from lower to higher trophic levels
Top-Down model of community organization
"trophic cascade model" proposes that control comes from the trophic level above; in this case predators control herbivores, which in turn control primary producers 
What determines community structure in bottom-up model?
presence or absence of mineral nutrients, inlcuding abundance of primary producers
Nonequilibrium Model
describes communities as constantly changins after being buffeted by disturbances
Disturbance
an event that changes a community, removes organisms from it and alters resource availability
Example of Disturbance
fire- in most terrestrial ecosystems
Intermediate Disturbance hypothesis
suggests the moderate levels of disturbance can foster greater diversity than either high or low levels of disturbance
Low levels of Disturbance
allows dominant species to exclude less competitive species
Two key factors that affect a community's species diversity
latitude and area
Species richness
declines along an equatorial-polar gradient and is especially great in the tropics
Climate
primary cause of the latitudinal gradient in biodiversity
Two climatic factors correlated with biodiversity
solar energy and water availability; can be considered together by measuring a community's rate of evapotranspiration
Evapotranspiration
evaporation of water from soil plus transpiration of water from plants
Species-area curve
quantifies the idea that, all other factors being equal, a larger geographic area has more species
Examples of species-area curve
studies of species richness on the Galapagos Islands support the prediction that species richness increases with island size; USA- breeding birds supports this
Pathogens
effects ecological communities and their structures; inclide disease-causing microorganisms, viruses, viroids and prions
What is transporting pathogens?
human activities at unprecedented rates 
Zoonotic
pathogens that have been transferred from other animals to humans
How can pathogens be transfered?
directly or through an intermediate species called a vector
Ecosystem
consists of all the organisms living in a community, as well as the abiotic facors with which they interact
Range of ecosystems
can range from a microcosm, such as an aquarium, to a large areas, such as a lake
Ecosystem dynamics involve two main processes
energy flow and chemical cycling
Energy and matter in respect to ecosystems
energy flows through while matter cycles within them
First law of thermodynamics
energy cannot be created or destroyed, only transformed
2nd law of thermodynamics
every exchange of energy increases the entropy of the universe
Energy flow through ecosystem
energy enters as solar radiation, is conserved, and is lost from organisms as heat; not always completely efficient
Law of conservation of mass
states that matter cannot be created or destroyed
Chemical elements and ecosystems
they are continually recycled through
Forest ecosystem (conservation of mass)
most nutrients enter as dust or solutes in rain and are carried away in water
Autotrophs
build molecules themselves using photosynthesis or chemosynthesis as an energy source
Heterotrophs
depend on the biosynthetic output of other organisms
Energy and Nutrient Flow
primary producers--primary consumers--secondary consumers--tertiary consumers
Primary Producers
autotrophs
Primary consumers
herbivores
Secondary consumers
carnivores
Tertiary consumers
carnivores that feed on other carnivores
Detritivores
"decomposers" are consumers taht derive their energy from detritus; ex: prokaryotes and fungi
Detritus
nonliving organic matter
What connects all trophic levels?
decomposition
Secondary production
amount of chemical energy in food converted to a new biomass during a given period of time
Trophic efficiency
percentage of production transferred from one trophic level to the next and it usually ranges from 5% to 20%
How efficient is the energy transfer between trophic levels?
10%
What percent of chemical energy fixed by photosynthesis reaches a teritary consumer?
0.10%
Biogeochemical cycles
nutrient circuits in ecosystems involve biotic and abiotic components and are called this
Element cycle
carbon, oxygen, sulfur and nitrogen occur in the atmosphere and cycle globally while elements sucj as phosphorus, potassium and calciumcycle on a more local level
Carbon reservoirs
include fossil fuels, soils and sediments, solutes in oceans, plant and animal biomass, and the atmosphere
Nitrogen is a component of...
amino acids, proteins, and nucleic acids
Main reservoir of nitrogen
atmosphere; through this it must be converted to NH4+ or NO3- for uptake by plants, via nitrogen fixation by bacteria
How is organic nitrogen decomposed?
decomposed to NH4+ by ammonification and NH4+ is decomposed to NO3- by nitrification
Dentrification and Nitrogen
converts NO3- back to N2
Rates at which nutrients cycle in different ecosystems
vary greatly, mostly as a result of differing rates of decomposition
Rate of decomposition
controlled by temperature, moisture, and nutrient availability
Agriculture and chemical cycles on Earth
removed from ecosystems nutrients that would ordinally be cycled back into the soil; nitrogen is main nutrient lost through agriculture
What is used to replace lost nitrogen through agriculture?
industrially produced fertilizer and effects on the ecosysyem can be harmful
Critical load
for a nutrient is the amount of plants can absorb without damaging the ecosystem
The critical load is exceeded when...
excess nutrients are added to an ecosystem
Eutrophication
excessive algal growth that can greatly harm freshwater ecosystems
Acid precipitation 
pH less than 5.6 
Cause of acid precipitation
combustion of fossil fuels
Why are toxins harmful to the environment?
become more concentrated in successive trophic levels
Biological Magnification
concentrates toxins at higher trohpic levels, where biomass is lower
What are subject to biological magnification in ecosystems?
PCBs (polychlorinated biphenyls) and many pesticides such as DDT
Greenhouse effect
CO2, water vapor, and other greenhouse gases reflect infared radiation back toward Earth
What could cause global warming and climate change?
increased levels of atmospheric CO2 are magnifying the greenhouse effect
Where shows the strongest effect of global warming?
northen coniferous forests and tundra
How can global warming be slowed?
reducing energy needs and converting to renewable sources of energy

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