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Viscosity
the thickness of a fluid that causes objects to encounter resistance as they move through it
Saturation
an upper limit of solubility in water
Solutes
dissolve substances in water
Semipermeable membranes
membranes that allow only particular molecules to pass through; reduces free movement of solutes
Osmosis
movement of water across a semi-permeable membrane
Osmotic potential
the force with which a solution attracts water by osmosis
Osmoregulation
mechanisms organisms use to maintain a proper solute balance
Boundary layer
a region of unstirred air or water that surrounds the surface of an object; removed gases from this region are slow to be replaced; this further limits carbon availability
Countercurrent circulation
an adaptation where blood and water flow in opposite directions so that the concentration of O2 in water is always greater than the concentration of blood
Q10 value
a ratio of a physiological process rate at one temperature to the rate of that process when the temperature is 10deg Celsius cooler
Thermal pollution
changing the temperature of an environment via human discharges ex: effluent from nuclear power plants
Glycerol and glycoproteins
chemicals present in some animals that prevent freezing by reducing strength of hydrogen bonds or via supercooling ex: coating of ice seeds
Thermal optima
the range of temperature in which an organism best performs
Isozymes
different forms of an enzyme that catalyze a reaction
Cohesion
the mutual attraction of water molecules; allows water to move up through empty remains of xylem cells
Root Pressure
when osmotic potential in the roots of a plant draws in the water from the soil and forces it into xylem
Transpiration
the process by which leaves can generate water potential as water evaporates from the surfaces of leaf cells
Cohesion-tension theory
the mechanism of water movement from roots to leaves due to water cohesion and water tension
Stomata
Small openings on leaf surfaces that are points of entry for CO2 and exit points for water vapor; bordered by guard cells that open and close each stoma
Electromagnetic radiation
energy from the sun; packaged in small, particle-like units called photons
Photosynthetically active region
wavelengths of light that are suitable for photosynthesis; includes wavelengths from 400nm(violet) to 700nm (red)
Chloroplasts
specialized cell organelles found in eukaryotic photosynthetic organisms
Photosynthesis
known as light reactions and the Calvin cycle; takes place in the stroma of chloroplasts; energy in ATP and NADPH is used to convert CO2 into glucose
C3 Photosynthesis
Process is catalyzed by RuBP carboxylase-oxidase CO2 + RuBP-->2G3P Common, Rubisco, inefficient
Disadvantages of C3 Photosynthesis
Rubisco is inefficient; low affinity for CO2 so plants need large amounts; Rubisco also preferentially binds to O2 which is problematic in hot and dry conditions ; Stromata close to prevent O2 from leaving the leaf
Photorespiration
reversal of light reactions to oxidate carbohydrates to CO2 and H2O by Rubisco
C4 Photosynthesis
a photosynthetic pathway in which CO2 is initially assimilated into a four-carbon compound, oxaloacetic acid(OAA); provides an advantage in hot and dry conditions CO2 + PEP --> OAA derived, PEP and OAA, reactions physically seperated
CAM Photosynthesis
a pathway in which the initial assimilation of carbon into OAA occurs at night; stromata closes during the say to reduce transpiration rates derived, PEP and OAA, temporal separation of reactions
Homeostasis
an organisms ability to maintain constant internal conditions int eh face of a varying external enviornment
Negative Feedback
the action of internal response mechanisms that restors a system to a desired state, or set point, when the system deviates from that state
Radiation
the emission of electromagnetic energy by a surface, increases with the fourth power of absolute temperature
Conduction
the transfer of the kinetic energy of heat between substances that are in contact with one another; rates depend on surface ares, resistance of substances to heat transfer, and temperature differences between substances
Convection
the transfer of heat by movement of liquids and gases; molecules next to a warm surface gain energy and move away
Evaporation
the transformation of water from a liquid to a gaseous state with the input of energy; removes heat from a surface
Thermal inertia
the resistance to a change in temperature due to a large body volume
Thermoregulation
the ability of an organism to control the temperature of its body
Homeotherms
organisms that maintain constant temperature; this allows biochemical reactions to work most efficiently
Poikilotherms
organisms that so not have constant body temperatures
Ectotherms
organisms with body temperatures determined by their external environment; not necessarily poikilotherms
Endotherms
organisms that can generate metabolic heat to raise body temperature higher than the external environment
Blood Shunting
when specific blood vessels shut off so less of an animal's warm blood flows to cold extremities where heat would be lost; occurs at pre-capillary sphincters
Deoxyribonucleic acid (DNA)
a molecule composed of two strands of nucleotides that are wound together into a double helix; each strand is composed of subunits called nucleotides; each nucleotide has a sugar, phosphate group, and one of the four nitrogenous bases (A,T,G,C)
Chromosomes
compact structures consisting of long strands of DNA wound around proteins
Alleles
different forms of a particular gene
Polygenic
when a single trait is affected by several genes; enables phenotypes to span a range of values in a population
Pleiotrophy
when a single gene affects multiple traits
Epistasis
when the expression of one gene is controlled by another gene ex: mouse hair color is determined by a gene that codes for black or brown pigments, a second gene determines whether the hair will have any pigment at all
Heterozygous
when an individual has two different alleles of a particular gene
Homozygous
when an individual has two identical alleles of a particular gene
Codominant
when two alleles both contribute to the phenotype ex: flower color in snapdragons
Dominant
an allele that masks the expression of the other allele
Recessive
an allele whose expression is masked by another allele; most harmful alleles are recessive
Gene Pool
collection of alleles from all individuals in a population
Random Assortment
the process of making haploid gametes in which the combination of alleles that are placed into a given gamete could be any combination of those possessed by the diploid parent
Mutation
a random change in the sequence of nucleotides in regions of DNA that controls the expression of a gene
Recombination
the reshuffling of genes that can occur as DNA is copied during meiosis and chromosomes exchange genetic material
Genetic Drift
a process that occurs when genetic variation is lost due to random variation in mating, mortality, fecundity, and inheritance
Bottleneck Effect
a reduction of genetic diversity in a poplutaion cue to large reduction in population size
Founder Effect
when a small number of individuals leave a large population to colonize a new area and bring with them only a small amount of genetic variation
Selection
the process by which certain phenotypes are favored to survive and reproduce over other phenotypes; relationship between phenotype and fitness
Stabilizing selection
when individuals with intermediate phenotypes have higher survival and reproductive success than those with extreme phenotypes
Directional selection
when individuals with extreme phenotypes experience higher fitness than the average population phenotype ex: drought on the Galapagos Island increased the proportion of large seeds; Birds with large beaks were better able to consume large seeds, so large beaks were selected for
Disruptive selection
when individuals withe either extreme phenotype experience higher fitness than individuals with an intermediate phenotype ex: Mexican spade-foot toad can have mouth morphology to be carnivorous, omnivorous, or intermediate consumer
Microevolution
the evolution of populations; affected by random processes and selection
Artificial selection
selection in which humans decide which individuals will breed; breeding is one with a preconceived goal for the traits in the population
Industrial melanism
a phenomenon in which industrial activities cause habitats to become darker due to population; individuals possessing darker phenotypes are favored by selection ex: during the 19th century, peppered mother were white with black spots
Macroevolution
evolution at higher levels or organization including species, genera, families, orders, and phyla
Speciation
the evolution of new species
Phylogenetic trees
hypothesized pattern of relatedness among different groups such as populations, species, or genera; depict which groups evolved from other groups
Allopatric Speciation
the evolution of new species through the process of geographic isolation
Sympatric speciations
the evolution of new species without geographic isolations
Polyploid
a species that contains three or more sets of chromosomes; can also give rise to sympatric speciation
Weather
the variation in temperature and precipitation over periods of hours or days
Climate
the typical atmospheric conditions that occur through the year, measured over many years
Large-scale spatial variation
may be caused by factors such as climate, land topography, and soil type
Small-scale spatial variation
is generated by factors such as plant structure and animal behavior
Phenotypic trade-off
a situation in which a given phenotype experiences higher fitness in one environment, whereas other phenotypes experience higher fitness in other environments
Phentotypic plasticity
the ability of a single genotype to produce multiple phenotypes; allows organisms to achieve homeostasis if environmental conditions vary
Acclimation
an environmentally induced change in an individual's physiology, can also be relatively rapid
hermaphrodites
individuals that produce both male and female gametes; individuals are able to fertilize their eggs with their own sperm
Inbreeding depression
the decrease in fitness caused by matings between close relatives due to offspring inheriting deleterious alleles from both the eggs and the sperm
Microhabitats
locations within a habitat that differ in environmental conditions from the rest of the habitats
Dormacy
a condition in which organisms dramatically reduce their metabolic process 4 types: Diapause, Hibernation, Torpor, Aestivation
Diapause
involves a partial or complete physiological shutdown in response to unfavorable conditions; common in insects
Hibernation
individuals reduce the energetic costs of being active by lowering heart rate and decreasing body temperature; common in mammals
Torpor
a brief period of dormancy in which individuals reduce activity and body temperature; common in birds and mammals
Aestivation
the shutting down of metabolic processes during the summer in response to hot or dry conditions; examples include snails, desert tortoises, and crocodiles
Central plane foraging
foraging behavior in which acquired food is brought to a central plaace
Risk-sensitive foraging
foraging behavior that is influenced by the presence of predator
Handling time
the amount of time that a predator takes to consume a captured prey
Life history
the schedule of an organism's growth, development, reproduction, and survival; represents an allocation of limited time and resources to achieve maximum reproductive success
Fecundity
the number of offspring produced by an organism per reproductive episode
Parity
the number of reproductive episodes an organism experiences
Parental investment
the time and energy given to an offspring by its parents
Longevity (life expectancy)
the life span of an organism
"Sloe" life history
long time to sexual maturity; long life spans; low numbers of offspring; high parental investment ex: elephants, oak trees
"Fast" life history
short time to sexual maturity; short life span; hight numbers of offspring; little parental investment ex: fruit flies, weeds
J. Philip Grime
proposed that plant life history depends on stress, competition, and the frequency of disturbances
Principle of Allocation
the observation that when resources are devoted to one body structure, physiological function, or behavior, they cannot be allotted to another
Determiate growth
a growth pattern in which an individual does not grow any more once it initiates reproduction; occurs in many species of birds and mammals
Indeterminate growth
a growth pattern in which an individual continues to grow after it initiates reproduction; occurs in many species of plants, invertebrates, fishes, reptiles, and amphibians
Semelparity
when organisms reproduce only once during their life; relatively rare in vertebrates, but common in insects or plants
Iteroparity
when organisms reproduce multiple times during their life; common among birds, reptiles, mammals, and amphibians
Annual
an organism that has a life span of one year
Perennial
an organism that has a life span of more than one year
Photperiod
the amount of light that occurs each day; provides a cue for many events in the life histories of virtually all organisms
Secual Reproduction
a reproduction mechanism in which progeny inherit DNA from two parents
Gonads
the primary sexual organs in animals
Asexual Reproduction
a reproduction mechanism in which progeny inherit DNA from a single parent
Vegetative reproduction
a form of asexual reproduction in which an individual is produced from the nonsexual tissues of a parent
Clones
individuals that descend asexually from the same parent and bear that same genotype
Parthenogenesis
a form of asexual reproduction in which an embryo is produced without fertilization
Purging mutations
sexually reproducing organisms can lose deleterious mutations during meiosis
Coping with environmental variation
offspring are likely to encounter different environmental conditions than their parents do
Red Queen Hypothesis
sexual selection allows hosts to evolve at a rate that counters the rapid evolution of parasites
Perfect flower
flowers that contain both male and female flowers
Simultaneous hermaphrodites
individuals that possess male and female reproductive functions at the same time
Sqeuential hermaphrodites
individuals that possess male or female reproductive function and then switch to the other
Monoecious
plants that have separate male and female flowers on the same individual
Dioecious
plants that contain either only male flowers or only female flowers on a single individual
Enviornmental sex determination
a process in which sex is determined largely by the environment; this is a type of phenotypic plasticity, where the phenotype is sex
Temperature-dependent sex determination
occurs when the sex of an individual is determined by the temperature at which eggs develop
Frequency dependent selection
when the rarer phenotype in a population is favored by natural selection
Local mate competition
when competition for mates occurs in a very limited area, and only a few males are required to fertilize all of the females
Mating System
the number of mates each individual has and the permanence of the relationship with those mates
Promiscuity
males mate with multiple females and females mate with multiple males and do not create lasting social bonds; common among animals and outcrossing plants
Polygamy
a single individual of one sex forms long-term social bonds with more than one individual of the opposite sex
Polygyny
a polygamous mating system in which makes with more then one female
Polyandry
a polygamous mating system in which a female mates with more than one male
Monogamy
when a social bond between a male and female persists through the period that is required for them to rear offspring
Extra-pair copulation
when an individual that has a social bond with a mate also breeds with other individuals
Mate Guarding
a behavior in which one partner prevents the other partner from participating in extra-pair copulations
Sexual selection
differential survival and reproduction due to sex-specific traits that are related to reproduction; leads to a variety of differences between male and females
Sexual dimorphism
the difference in the phenotype between males and females of the same speices
Primary Sexual Characterists
traits related to fertilization
Seconday SexualCharacteristics
traits related to differences between the sexes in terms of body size, ornaments, color, and courtship
Fecundity-body size ratio
body size differences between sexes are common in animals; there has been selection for an increased number of gametes or an increase in parental care in one of the sexes
Contest
when males compete for females, selection will favor the evolution of weapons
Mate Choice
female choice
Good gGenes Hypothesis
the hypothesis that an individual chooses a mate that possesses a superior genotype
Good Health Hypothesis
the hypothesis that an individual chooses the healthiest mates
Runaway Sexual Selection
when selection for preference of a sexual trait and selection for that trait continue to reinforce each other; continues until males run out of genetic variation
The Handicap Principle
the greater the handicap an individual carries, the greater its ability must be to offset that trait
Social Behavior
interactions with members of one's own species, including mates, offspring, other relatives, and unrelated individuals
Predator Dtetection/Evasion
a group may be able to fend off predatos better than an individual
Dilution Effect
the reduced, or diluted, probability or predation to a single animal when it is in a group; probability of death=1/group size
Vigilance-Group Size Effect
more individuals watching the predators allows each individual to spend less time watching, and more time feeding
Food Loaction
many individuals searching for food may be able to find rare food more easily
Mate finding
being social makes it easier to find potential mates because large groups attract the attention of females
Lek
the location of an animal aggression to put on display to sttract the opposite sex
Conpicuousness
group of animals are more conspicuous to predators
Disease Transmission
the rise of parasites increases in groups; high densities can increase the rate at which diseases spread
Competition
larger groups are better able to locate food, but that food must be shared among all members
Aggression
living in groups can lead to aggression among members
Territory
any area defended by one or more individuals against the intrusion of others
Dominance hierarchy
a social ranking among individuals in a group typically determined through contests such as fighting or other contests of strength or skill
Game-Theory Model
model that attempts to predict the outcome of behavioral decisions when those decisions depend on behavior of other players
Donor
the individual who directs a behavior toward another individual as part of a social interaction
Recipient
the individual who receives the behavior of a donor in a social interaction
Cooperation
when the donor and the recipient of a social behavior both experience increased fitnedd from an interaction
Selfishness
when the donor of a social behavior experiences increased fitness and the recipient experience decreased fitness
Spitefulness
when a social interaction reduces the fitness ofboth donor and recipient
Alturigm
a social interaction that increases recipient fitness and decreases the fitness of the donor
Indirect fitness
the fitness that an individual gains by helping relatives pass on copies of their genes
Inclusive fitness
the sum of direct and indirect fitness
Direct Fitness
the fitness an individual gains by passing on copies of its genes to its offspring
Die=rect selection
selection that favors direct fitness
Indirect selection (kin selection)
selection favoring indirect fitness
Eusocialanimals
1.Several adults living together 2.Overlapping generations of parents and offspring living together in the same group 3. Cooperation in nest building and brood care 4. Reproductive dominance by one or a few individuals, and the presence of sterile individuals
Caste
individuals within a social group sharing a specialized form of behavior
Queen
the dominant, egg-laying female in eusocial insect societies; typically mate once during their lives
Haplodiploid
a sex-determination system in which one sex is haploid and other sex is diploid
Evolution of Eusociality
has independently evolved many times; being haplodiploid favors the evolution eusociality; could evolve if the cost of leaving a colony is high due to a low likelihood of surviving, this reduces the cost of foregoing reproduction

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