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Lardizabalaceae

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LARDIZABALACEAE1. DECAISNEA J. D. Hooker & Thomson, Proc. Linn. Soc. London 2: 350. 1855, nom. cons.1. Decaisnea insignis (Griffith) J. D. Hooker & Thomson, Proc. Linn. Soc. London 2: 349. 1855.2. AKEBIA Decaisne, Compt. Rend. Hebd. Séances A1. Akebia quinata (Houttuyn) Decaisne, Arch. Mus. Hist. Nat. 1: 195. 1839.2. Akebia longeracemosa Matsumura, Bot. Mag. (Tokyo) 13: 18. 1899.3. Akebia trifoliata (Thunberg) Koidzumi, Bot. Mag. (Tokyo) 39: 310. 1925.3a. Akebia trifoliata subsp. trifoliata3b. Akebia trifoliata subsp. australis (Diels) T. Shimizu, Quart. J. Taiwan Mus. 14: 201. 1961.3c. Akebia trifoliata subsp. longisepala H. N. Qi4. Akebia chingshuiensis T. Shimizu, Quart. J. Taiwan Mus. 14: 201. 1961.3. ARCHAKEBIA C. Y. Wu, T. Chen & H. N. Qin in H. N. Qin, Acta Phytotax. Sin. 33: 240. 1995.1. Archakebia apetala (Q. Xia, J. Z. Suen & Z. X. Peng) C. Y. Wu, T. Chen & H. N. Qin in H. N. Qin, Acta Phytotax. Sin. 33: 240. 1995.4. HOLBOELLIA Wallich, Tent. Fl. Napal. 1: 23. 1824.1. Holboellia coriacea Diels, Bot. Jahrb. Syst. 29: 342. 1900.2. Holboellia parviflora (Hemsley) Gagnepain, Bull. Mus. Hist. Nat. (Paris) 14: 68. 1908.3. Holboellia brachyandra H. N. Qin, Cathaya 8–9:4. Holboellia grandiflora Réaubourg, Bull. Soc. 5. Holboellia angustifolia Wallich, Tent. Fl. Napal 1: 25. 1824.5a. Holboellia angustifolia subsp. angustifolia5b. Holboellia angustifolia subsp. linearifolia T5c. Holboellia angustifolia subsp. trifoliata H. 5d. Holboellia angustifolia subsp. obtusa \(Gagn6. Holboellia latifolia Wallich, Tent. Fl. Napal. 1: 24. 1824.6a. Holboellia latifolia subsp. latifolia6b. Holboellia latifolia subsp. chartacea C. Y. W7. Holboellia medogensis H. N. Qin, Cathaya 8–9: 8. Holboellia chapaensis Gagnepain, Bull. Soc. Bot. France 85: 165. 1938.9. Holboellia pterocaulis T. Chen & Q. H. Chen, Fl. Guizhou 2: 675. 147. 1986.5. STAUNTONIA de Candolle, Syst. Nat. 1: 511, 513. 1817.1. Stauntonia decora (Dunn) C. Y. Wu ex S. H. Huang, Fl. Yunnan. 2: 8. 1979.2. Stauntonia brunoniana Wallich ex Hemsley, Hook3. Stauntonia elliptica Hemsley, Hooker’s Icon. P4. Stauntonia maculata Merrill, Lingnan Sci. J. 13: 24. 1934.5. Stauntonia chinensis de Candolle, Syst. Nat. 1: 514. 1817.6. Stauntonia glauca Merrill & Metcalf, Lingnan Sci. J. 16: 80. 1937.7. Stauntonia purpurea Y. C. Liu & F. Y. Lu, Quart. J. Chin. Forest. 11: 110. 1978.8. Stauntonia trinervia Merrill, Lingnan Sci. J. 13: 24. 1934.9. Stauntonia duclouxii Gagnepain, Bull. Soc. Bot. France 55: 48. 1908.10. Stauntonia libera H. N. Qin, Cathaya 8–9: 13611. Stauntonia oligophylla Merrill & Chun, Sunyatsenia 5: 54. 1940.12. Stauntonia obcordatilimba C. Y. Wu & S. H. Huang in S. H. Huang, Fl. Yunnan. 2: 11. 1979.13. Stauntonia yaoshanensis F. N. Wei & S. L. Mo, Guihaia 3: 309. 1983.14. Stauntonia conspicua R. H. Chang, Acta Phytotax. Sin. 25: 235. 198715. Stauntonia pseudomaculata C. Y. Wu & S. H. Huang in S. H. Huang, Fl. Yunnan. 2: 11. 1979.16. Stauntonia obovatifoliola Hayata, Icon. Pl. Formos. 8: 4. 1919.16a. Stauntonia obovatifoliola subsp. obovatifoliola16b. Stauntonia obovatifoliola subsp. urophylla 17. Stauntonia cavalerieana Gagnepain, Bull. Soc. Bot. France 55: 47. 1908.18. Stauntonia obovata Hemsley, Hooker’s Icon. Pl19. Stauntonia brachyanthera Handel-Mazzetti, Anz. Akad. Wiss. Wien, Math.-Naturwiss. Kl. 58: 90. 1921.20. Stauntonia leucantha Diels ex Y. C. Wu, Notizbl. Bot. Gart. Berlin-Dahlem 13: 373. 1936.6. SINOFRANCHETIA \(Diels\) Hemsley, Hooker’s 1. Sinofranchetia chinensis \(Franchet\) Hemsl7. SARGENTODOXA Rehder & E. H. Wilson in Sargent, Pl. Wilson. 1: 350. 1913.1. Sargentodoxa cuneata (Oliver) Rehder & E. H. Wilson in Sargent, Pl. Wilson. 1: 351. 1913.Flora of China 6: 440–454. 2001. LARDIZABALACEAE 木通科 mu tong ke Chen Dezhao (陈德昭 Chen Te-chao)1; Tatemi Shimizu2 Woody climbers, rarely erect shrubs (Decaisnea). Monoecious, rarely dioecious. Stems climbing or twining (erect in Decaisnea); wood with broad medullary rays. Winter buds large; outer scales 2 to numerous; scales subtending leaves persistent, scaly. Stipules usually absent (present and very small in Sinofranchetia). Leaves alternate, palmate or 3-foliolate, rarely pinnate (Decaisnea). Petiole and petiolules swollen at both ends, articulate. Flowers functionally unisexual by reduction or abortion, actinomorphic. Sepals 6 (usually 3 in Akebia), petaloid, in 2 whorls, imbricate or outer 3 valvate. Petals 6, nectariferous, much smaller than sepals or absent. Male flowers: stamens free or ± connate into a tube; anthers 2-celled, extrorsely dehiscent by longitudinal slit; connective apical appendage hornlike or apiculate, sometimes unappendaged. Pistillodes 3–6(–9), small, filiform. Female flowers: staminodes 6. Carpels 3(–9) or numerous (Sargentodoxa), superior, free; ovules numerous, orthotropous or anatropous, in 2 to several longitudinal rows on laminar placenta, rarely 1, pendulous, hemitropous to subanatropous (Sargentodoxa). Stigma conspicuous, sessile or subsessile. Fruit fleshy follicles or baccate, indehiscent or dehiscent along abaxial suture. Seeds numerous, rarely solitary; seed coat crustaceous; endosperm copious; embryo small, straight. Nine genera and ca. 50 species: mainly in E Asia, two monotypic genera in S America; seven genera (two endemic) and 37 species (25 endemic) in China. Chen Te-chao. 2001. Lardizabalaceae. In: Ying Tsun-shen, ed., Fl. Reipubl. Popularis Sin. 29: 1–50. 1a. Stems erect; leaves odd pinnate with 13–25 leaflets; inflorescences terminal; winter bud outer scales 2 1. Decaisnea 1b. Stems climbing or twining; leaves palmate or (1 or)3–9-foliolate; inflorescences axillary; winter bud outer scales numerous. 2a. Leaves (1 or)3-foliolate; lateral leaflets asymmetric. 3a. Stem without red sap when cut; lateral leaflets slightly asymmetric at base, smaller than terminal leaflet; carpels 3 in female flowers, ovules ca. 20 per carpel; mature carpels 1–3 at each node of infructescence ................................................................................................................... 6. Sinofranchetia 3b. Stems with red sap when cut; lateral leaflets extremely asymmetric on one side, larger than terminal leaflet; carpels numerous in female flowers; ovules 1 per carpel; mature carpels 20–40 spirally arranged on an enlarged receptacle ................................................................................................... 7. Sargentodoxa 2b. Leaves palmately compound,


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