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Quaternary Research 67 2007 143 151 www elsevier com locate yqres Dung diet and the paleoenvironment of the extinct shrub ox Euceratherium collinum on the Colorado Plateau USA Manny Kropf a Jim I Mead a c R Scott Anderson a b a b Laboratory of Quaternary Paleontology Quaternary Sciences Program Northern Arizona University Flagstaff AZ 86011 USA Center for Environmental Sciences and Education and Quaternary Sciences Program Northern Arizona University Flagstaff AZ 86011 USA c Department of Geology Northern Arizona University Flagstaff AZ 86011 USA Received 17 May 2006 Available online 7 November 2006 Abstract Fossil remains of Euceratherium collinum extinct shrub ox have been found throughout North America including the Grand Canyon Recent finds from the Escalante River Basin in southern Utah further extend the animal s range into the heart of the Colorado Plateau E collinum teeth and a metapodial condyle foot bone have been recovered in association with large distinctively shaped dung pellets a morphology similar to a Hershey s Kiss HK from a late Pleistocene dung layer in Bechan Cave HK dung pellets have also been recovered from other alcoves in the Escalante River Basin including Willow and Fortymile canyons Detailed analyses of the HK pellets confirmed them to be E collinum and indicate a browser type diet dominated 95 by trees and shrubs Artemisia tridentata big sagebrush Acacia sp acacia Quercus oak and Chrysothamnus rabbit brush The retrieval of spring and fall pollen suggests E collinum was a year round resident in the Escalante River Basin 2006 University of Washington All rights reserved Keywords Pleistocene Dung Bechan Cave Escalante River Basin Euceratherium Paleoenvironment Microhistology Pollen Introduction The shrub ox Euceratherium collinum Artiodactyla Ruminantia Bovidae Ovibovini along with several related taxa represent a little studied and poorly understood group of extinct North American ovibovids Frick 1937 Scott 1962 Martin and Wright 1967



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