Coelomates: Protostomes and DeuterostomesProtostomes have spiral cleavage, the blastopore becomes themouth, mosaic development - mollusks, annelids, arthropodsDeuterostomes have radial cleavage, the blastopore becomes theanus, regulative development - echinoderms, chordatesspiral cleavageradial cleavageProtostomesDeuterostomesPhylum Echinodermata - Echinodermsincludes starfish, sea urchins,sand dollars, sea cucumbersName means “spiny skin” - havehard endoskeleton, often with spinesbelow thin skinEndoskeleton is made of calcium carbonate plates that form internallyPossess a unique water vascular system - a hydraulic system to aid movement and circulationMouth opens ventrally in most - anus is dorsalAdults have no head orbrainNervous system composed of circularnerve ring and radialbranchesSexes are separateFertilization is externalGonads (ovaries and testes) found in armsEchinoderms exhibit radial symmetry as adults - but have a bilaterally symmetric larvaTransform to pentaradial symmetry as they matureRadial symmetry works well for sessile organisms - primitive echinoderms (likesea lilies) are sessileBilateral larva thought to indicate that they originatedfrom a bilaterally symmetricancestor - original ancestor is uncertainThe Water Vascular SystemFive radial canals extend from a ring canal around the esophagusWater enters through madreporite, a sieve-like plateFlows to ring canal through the tubular stone canalRadial canals have lateral canals to tube-feetEach tube-foot has a muscular fluid-filled ampulla at its baseOperation of the water vascular systemAmpulla contracts, one way valve causes fluid to be forced into tube-foot - results in extensionFoot can attach to substrate, sometimes with suction-cup tipLongitudinal muscles contract and shorten tube-footWater forced back into ampullaRepeated movement results in locomotion or manipulation of objectsHave large coelom, lined with ciliaHelps provide for circulation and respirationIn most, respiration and waste removaloccur through skin gills (papulae) that project through skinEndoskeleton composed of ossiclesMovable or fixed plates under skinTube feet extend through perforationsReproduction - many species capable of extensive regenerationmay shed body parts when attacked -some reproduce asexually by splitting into partsSexual reproduction and fertilization is externalSexes are separate, have free-swimming bilaterally symmetric larvaThere are Six Living Classes of EchinodermsCrinoidea - sea lilies and feather starsAsteroidea - sea starsOphiuroidea - brittle starsEchinoidea - sea urchins and sand dollarsHolothuroidea - sea cucumbersConcentricycloidea - sea daisies - recently discoveredClass Crinoidea: The Sea Lilies and Feather StarsSessile, sedentary animalsMouth and anus located on upper surface of an open diskMany highly branched arms located around central disk - 5 to 200arms - smaller pinnules branch from each armfood collected by mucus on pinnules - transported by tube-feetFossils abundant, many more extinct forms than living speciesSea lilieshave a stalkFound inwater deeperthan 100 mClass Asteroidea - Sea Starsthe familiar "starfish"Active, important marine predatorsAbundant in shallow and deep watersArms merge gradually with diskArms in multiples of five, usually fiveBody flattened, flexible, covered with pigmented epidermisOften feed on bivalvesGrasp shell with tube feetto wrench openExtrude stomach into shellSecretes enzymes, digestssoft parts of bivalveMany are scavengersClass Ophiuroidea - Brittle StarsLargest class of echinodermsCalled “brittle” because they shed theirarms readilyFound in shallow water and one of most abundant animals in the deep-seaSlender arms set off sharply from central diskActively move arms for locomotionArms may be covered with spinesSome use arms to swimTube feet are important sensory organsTube feet capture small food particlesand move food to mouthClass Echinoidea - Sea Urchins and Sand DollarsLack distinct arms, but still have five-part body planFive rows of tube feet protrude from plates of skeletonEndoskeleton is made of fused platesSea urchins are globular with large spinesSand dollars are flattenedWalk with tube feet ormovement of spinesFeed on algaescraped off thesurfaces bytriangular teeth orfeed on food itemsfound on or in thesubstrateClass Holothuroidea - Sea CucumbersSoft, sluglike organisms with tough, leathery skinBilaterally symmetric externally, with five-part internal body planMouth is located on one end, anus on otherMouth surrounded by tube feet modified into tentaclesTentacles secrete mucus to trap food particlesEndoskeleton reduced to widely separated,microscopic platesRespiration by respiratory tree that originates at anusHave tube feet on bodyMove by tube feet or flexing of entire bodyMost are sluggishMay eject digestive system or respiratory tree when threatened by a